Professor Mark Willems
Professor of Exercise Physiology
Mark started to work at the University of Chichester in 2003. He was a member of the National Dutch Youth Handball Team and the National Dutch Military Team and won National Championships with handball at the junior and senior level in the Netherlands. Mark enjoys running at a speed at which you can enjoy the countryside.
- 1990 – MSc in Exercise Physiology, Vrije Universiteit Amsterdam, The Netherlands
- 1990 – MSc in Functional Anatomy, Vrije Universiteit Amsterdam, The Netherlands
- 1994 – PhD in Human Movement Sciences – Architectural Heterogeneity and Function of Bi-articular Muscle. Vrije Universiteit Amsterdam, The Netherlands
Mark is programme coordinator for the MSc in Sport and Exercise Physiology and has responsibility for the following modules:
- ESP504: Sports Science: Physicality of Gaming
Aims: The module aims to develop knowledge on the nature of the physical demands in response to active involvement in playing esports. This module will address the physical demands and preparation for esports performance with consideration of the role of the central nervous system, energy demands, nutrition, repetitive movements and endurance activities. Athletic esports demands and training will be discussed with focus on the metabolic, neuromuscular, neuroendocrine and cardiovascular systems of the human body to develop knowledge on the nature of esports performance.
- IOS7103: Muscle Physiology for Sport, Exercise and Nutrition
Aims: The module aims to develop a critical knowledge of muscle functioning, the nature of metabolic limitations in skeletal muscles during exercise, the role and function of skeletal muscles in specific sports, and the relationships between metabolic limitation, nutritional interventions, and training adaptations. The application of this knowledge to athletic training in divergent sporting activities, and general conditioning in sedentary and special populations will be discussed within both theoretical and practical contexts.
- IOS7015: Planning your Research Dissertation/Supervised Experience Portfolio
Aims: The module will aim to support the student in 1) the development of the research attitude and mindset capable of undertaking and completing a Research Dissertation or Supervised Portfolio, 2) the development and application of the specific research skills and measurements, and 3) the completion of the required paperwork for obtaining approval for initiation of the Research Dissertation or Supervised Experience Portfolio.
Mark has research interests in the following areas: Muscle Physiology, Muscle Fatigue, Muscle Injury, Ageing and Exercise, Eccentric Exercise, Occupational Physiology, Supplementation and Exercise (i.e. New Zealand blackcurrant and Matcha green tea), Exercise in Clinical Populations.
Director of Studies
Director of Studies
- 2006-2009 PhD: Sam Blacker – Physiological Effects of Prolonged Load Carriage by Backpack
- 2007-2010 PhD: Mandy Gault – Effects of Regular Downhill Walking on Cardiovascular Health and Physical Performance in Older People
- 2010-2013 PhD: James Gavin – The Effect of Glycogen Reduction on Neuromuscular Responses and Recovery from Eccentric Exercise
- 2013-2017 PhD: Matthew Cook – Effect of New Zealand Blackcurrant on Physiological and Metabolic Responses at Rest and During Exercise
- 2015-present MPhil/PhD: Ian Perkins – New Zealand Blackcurrant and Individual Responses to Exercise
- 2017-present MPhil/PhD: Stefano Montanari – Optimising Polyphenol Supplementation for Exercise Responses and Performance
European College of Sport Science (Fellow)
Lee BJ, Flood TR, Hiles AM, Walker EF, Wheeler LEV, Ashdown KM, Willems MET, Costello R, Greisler LD, Romano PA, Hill GW, Kuennen MR. Anthocyanin-rich blackcurrant extract preserves gastrointestinal barrier permeability and reduces enterocyte damage but has no effect on microbial translocation and inflammation after exertional heat stress. Int J Sport Nutr Exerc Metab. doi: 10.1123/ijsnem.2021-0330
Şahin MA, Bilgiç P, Montanari S, Willems MET. Intake Duration of Anthocyanin-Rich New Zealand Blackcurrant Extract Affects Cardiovascular Responses during Moderate-Intensity Walking But Not at Rest. .J Diet Suppl. doi: 10.1080/19390211.2021.2005214.
Cook MD, Dunne A, Bosworth M, Willems MET. Effect of Intake Duration of Anthocyanin-Rich New Zealand Blackcurrant Extract on Cardiovascular Responses and Femoral Artery Diameter during Sustained Submaximal Isometric Contraction. J Diet Suppl. doi: 10.1080/19390211.2021.1948943.
Costello R, Keane KM, Lee BJ, Willems MET, Myers SD, Myers F, Lewis NA, Blacker SD. Plasma uptake of selected phenolic acids following New Zealand blackcurrant extract supplementation in humans. J Diet Suppl. doi: 10.1080/19390211.2021.1914802.
Willems MET, Todaka M, Banic M, Cook MD, Sekine Y. Intake of New Zealand Blackcurrant Powder Affects Skin-Borne Volatile Organic Compounds in Middle-Aged and Older Adults. J Diet Suppl. doi: 10.1080/19390211.2021.1908479.
Willems MET. Anthocyanin-Rich New Zealand blackcurrant: Implications for Health. Agro FOOD Industry Hi Tech – vol31, 2022.
Willems MET, Blacker SD. Anthocyanin-Rich Supplementation: Emerging Evidence of Strong Potential for Sport and Exercise Nutrition. Frontiers in Nutrition 9, 864323, 2022. https://doi.org/10.3389/fnut.2022.864323.
Willems, MET, Banic M, Cadden R, Barnett L (2022) Enhanced walking-induced fat oxidation by New Zealand blackcurrant extract is body composition-dependent in recreationally active adult females. Nutrients 14(7), 1475. https://doi.org/10.3390/nu14071475.
Şahin MA, Bilgiç P, Montanari S, Willems MET. Intake Duration of Anthocyanin-Rich New Zealand Blackcurrant Extract Affects Metabolic Responses during Moderate Intensity Walking Exercise in Adult Males. J Diet Suppl. (in press). doi: 10.1080/19390211.2020.1783421.
Willems MET, Fry HL, Belding MA, Kaviani M. Three weeks daily intake of Matcha green tea powder affects substrate oxidation during moderate-intensity exercise in females. Journal of Dietary Supplements 18(5):566-576, 2021. doi: 10.1080/19390211.2020.1811443.
Fryer S, Giles D, Bird E, Stone K, Paterson C, Baláš J, Willems MET, Potter JA, Perkins IC. New Zealand blackcurrant extract enhances muscle oxygenation during repeated intermittent forearm muscle contractions in advanced and elite rock climbers. Eur J Sport Sci. 21(9):1290-1298, 2021. doi: 10.1080/17461391.2020.1827048.
Hossein M, Zahra A, Erfan B, Amir S, Sajjad R, Raúl D, Reid R, Emerson F, Mohammad HS, Majid SK, Mark E.T Willems, Ramin R, Alireza N. Acute doses of beetroot juice supplementation do not improve aerobic and anaerobic performance of trained male taekwondo athletes. International Journal of Environmental Research and Public Health, 18, 10202, 2021. doi:10.3390/ijerph181910202.
Fryer S, Paterson C, Perkins IC, Gloster C, Willems MET, Potter JA. New Zealand Blackcurrant Extract Enhances Muscle Oxygenation During Forearm Exercise in Intermediate-Level Rock Climbers. Int J Sport Nutr Exerc Metab. 30(4):258-263, 2020. doi: 10.1123/ijsnem.2019-0365.
Hiles AM, Flood TR, Lee BJ, Wheeler LEV, Costello R, Walker EF, Ashdown K, Kuennen MR, Willems MET. Dietary supplementation with New Zealand blackcurrant extract enhances fat oxidation during submaximal exercise in the heat. Journal of Science and Medicine in Sport 23(10):908-912, 2020.
Okamoto T, Hashimoto Y, Kobayashi R, Nakazato K, Willems MET. Effects of blackcurrant extract on arterial functions in older adults: A randomized, double-blind, placebo-controlled, crossover trial. Clin Exp Hypertens. 42(7):640-647, 2020. doi: 10.1080/10641963.2020.1764015.
Costello R, Willems MET, Myers SD, Myers F, Lewis NA, Lee BJ, Blacker SD No Effect of New Zealand Blackcurrant Extract on Recovery of Muscle Damage Following Running a Half-Marathon. Int J Sport Nutr Exerc Metab. 2020 May 29:1-8. doi: 10.1123/ijsnem.2019-0312.
Potter JA, Hodgson CI, Broadhurst M, Howell L, Gilbert J, Willems MET, Perkins IC. Effects of New Zealand blackcurrant extract on sports climbing performance. European Journal of Applied Physiology 120(1), 67-75, 2020.
Negaresh R, Del Coso J, Mokhtarzade M, Lima-Silva AE, Baker JS, Willems MET, Talebvand S, Khodadoost M, Farhani F. Effects of different dosages of caffeine administration on wrestling performance during a simulated tournament European Journal of Sport Science 19(4), 499-507, 2019. DOI: 10.1080/17461391.2018.1534990.
Willems MET, Şahin MA, Berendsen T, Cook MD. Effect of New Zealand blackcurrant on cycling performance and substrate oxidation in normobaric hypoxia in trained cyclists. Sports 7(3). pii: E67, 2019. doi: 10.3390/sports7030067.
Cook MD, Willems MET. Dietary Anthocyanins: A Review of the Exercise Performance Effects and Related Physiological Responses. International Journal of Sports Nutrition and Exercise Metabolism 29(3):322-330, 2019. doi: 10.1123/ijsnem.2018-0088.
Willems MET. Can you enhance exercise-induced fat oxidation with green tea drinking? Agro FOOD Industry Hi Tech – vol. 29(4) July/August 2018.
Willems MET, Parktin N, Widjaja W, Ajjimaporn A. Effect of New Zealand blackcurrant extract on physiological responses at rest and during brisk walking in Southeast Asian men: A randomized, double-blind, placebo-controlled, cross-over study. Nutrients 10(11). pii: E1732. doi: 10.3390/nu10111732.
Willems MET, Sahin MA, Cook MD. Matcha green tea drinks enhance fat oxidation during brisk walking in females. International Journal of Sports Nutrition and Exercise Metabolism 28(5):536-541, 2018. doi: 10.1123/ijsnem.2017-0237
Strauss JA, Willems MET, Shepherd SO. New Zealand blackcurrant extract enhances fat oxidation during prolonged cycling in endurance-trained females. European Journal of Applied Physiology 118(6), 1265-1272, 2018. doi: 10.1007/s00421-018-3858-3.
Kaya F, Yuktasir B, Bicer B, Willems MET, Yıldız N. The Effects of Two Different Stretching Programs on Balance Control and Motor Neuron Excitability. Journal of Education and Training Studies 6(5), 85-91, 2018.
Gavin JP, Myers SD, Willems MET. Cardiorespiratory and metabolic responses after exercise-induced muscle damage: the influence of lowered glycogen. The Journal of Sports Medicine and Physical Fitness 58(3), 332-340, 2018. doi: 10.23736/S0022-4707.16.06644-5.
Godwin C, Cook MD, Willems MET. Effect of New Zealand blackcurrant extract on performance during the running based anaerobic sprint test in trained youth and recreationally active male football players. Sports 5, 69, 2017. doi:10.3390/sports5030069.
Willems MET, Dos Santos Silva J, Cook MD, Blacker SD. Beneficial effects on fasting insulin and postprandial responses through 7-day intake of New Zealand blackcurrant powder. Journal of Functional Foods in Health and Disease 7(7), 483-493, 2017.
Gault ML, Willems MET. The metabolic equivalents of 1-mile walking by older adults; implications for health promotion. Health Promotion Perspectives 7(4), 216-222, doi: 10.15171/hpp.2017.38.
Cook MD, Myers SD, Gault ML, Willems MET. Blackcurrant Alters Physiological Responses and Femoral Artery Diameter during Sustained Isometric Contraction. Nutrients. 2017, 9(6). pii: E556. doi: 10.3390/nu9060556.
Murphy C, Cook MD, Willems MET. Effect of New Zealand blackcurrant extract on repeated cycling time trial performance. Sports 5(2), 25, 2017. doi: 10.3390/sports5020025.
Cook MD, Myers SD, Gault ML, Edwards VC, Willems ME. Dose effects of New Zealand blackcurrant on substrate oxidation and physiological responses during prolonged cycling. European Journal of Applied Physiology 117(6), 1207-1216, 2017. doi: 10.1007/s00421-017-3607-z.
Cook MD, Myers SD, Gault ML, Edwards VE, Willems MET. Cardiovascular Function during Supine Rest in Endurance Trained Males with New Zealand Blackcurrant: A Dose-Response Study. European Journal of Applied Physiology 117(2):247-254, 2017. doi: 10.1007/s00421-016-3512-x.
Naderi A, Sadeghi M, Sarshin A, Imanipour V, Nazeri SA, Farkhaie F, Willems MET. Muscle carnosine concentration with the co-ingestion of carbohydrate with β-alanine in male rats. Journal of Dietary Supplements 14(4), 373-379, 2017.
Ozmen T, Gunes GY, Dogan H, Ucar I, Willems M. The effect of kinesio taping versus stretching techniques on muscle soreness, and flexibility during recovery from Nordic hamstring exercise. Journal of body work and movement therapies 21(1), 41-47, 2017.
Naderi A, Earnest CP, Lowery RP, Wilson JM, Willems MET. Co-ingestion of nutritional ergogenic aids and high intensity exercise performance. Sports Medicine 46(10), 1407-1418, 2016. doi: 10.1007/s40279-016-0525-x.
Willems MET, Cousins L, Williams D, Blacker SD. Beneficial Effects of New Zealand Blackcurrant Extract on Maximal Sprint Speed during the Loughborough Intermittent Shuttle Test. Sports 4(3), 42, 2016. doi:10.3390/sports4030042.
Gavin JP, Myers SD, Mark E.T Willems. Effect of eccentric exercise with reduced muscle glycogen on plasma interleukin-6 and neuromuscular responses of m. quadriceps femoris. Journal of Applied Physiology (1985), 121(1):173-84, 2016. doi: 10.1152/japplphysiol.00383.2015.
Kulaksız TN, Koşar SN, Bulut S, Güzel Y, Willems MET, Hazir T, Turnagöl HH. Mouth Rinsing with Maltodextrin Solutions Fails to Improve Time Trial Endurance Cycling Performance in Recreational Athletes. Nutrients 8(5), E269, 2016. doi: 10.3390/nu8050269.
Naderi A, Far AH, Willems MET, Sadeghi M. Effect of Four Weeks of β-alanine supplementation on muscle carnosine and blood serum lactate during exercise in male rats. Journal of Dietary Supplements 13(5), 487-494, 2016. doi: 10.3109/19390211.2015.1113223.
Sozbir K, Willems MET, Tiryaki-Sonmez G, Ragauska P. Acute effects of contract-relax PNF and static stretching on flexibility, jump performance and EMG activities: A case study. Biology of Exercise 12(1), 33-55, 2016. doi: 10.4127/jbe.2016.0099.
Ozmen T, Aydogmus M, Dogan H, Acar D, Zoroglu T & Willems M. The effect of kinesio taping® on muscle pain, sprint performance and flexibility in recovery from squat exercise in young adult women. Journal of Sport Rehabilitation 25(1), 7-12, 2016. doi: 10.1123/jsr.2014-0243.
Gavin JP, Myers SD, Willems MET. The accumulative effect of concentric-biased and eccentric-biased exercise on cardiorespiratory and metabolic responses to subsequent low-intensity exercise: a preliminary study. Journal of Human Kinetics 49, 131-40, 2015 doi: 10.1515/hukin-2015-0115.
Cook MD, Myers SD, Blacker SD, Willems MET. New Zealand Blackcurrant Extract Improves Cycling Performance and Fat Oxidation in Cyclists. European Journal of Applied Physiology 115(11), 2357-2365, 2015.
Perkins IC, Vine SA, Blacker SD, Willems MET. New Zealand Blackcurrant Extract Improves High-intensity Intermittent Running. International Journal of Sports Nutrition and Exercise Metabolism 25, 487-493, 2015.
McGuire S, Willems MET. Physiological responses during multiplay exergaming in young adult males are game-dependent. Journal of Human Kinetics 46, 263-271, 2015.
Cook MD, Myers SD, Kelly JSM, Willems MET. Effect of Eccentric and Concentric Exercise on Breathing Efficiency. Sports 3, 12-20, 2015.
Willems MET, Myers SD, Gault ML & Cook MD. Beneficial physiological effects with blackcurrant intake in endurance athletes. International Journal of Sport Nutrition & Exercise Metabolism 25(4), 367-374, 2015.
Gavin JP, Myers SD & Willems MET. The effect of glycogen reduction on cardiorespiratory and metabolic responses during downhill running. European Journal of Applied Physiology 115(5), 1125-1133, 2015.
Gavin JP, Myers SD & Willems MET. Neuromuscular responses to mild-muscle damaging eccentric exercise in a low glycogen state. Journal of Electromyography and Kinesiology 25, 53-60, 2015.
Gallagher CA, Lewis MP, Willems MET & Myers SD. The application of maximal heart rate predictive equations in hypoxia 115(2), 277-284, 2015.
Gallagher CA, Willems MET, Lewis, MP & Myers SD. Effect of acute normobaric hypoxia on the ventilatory threshold. European Journal of Applied Physiology 114(8):1555-62, 2014.
Cook MD, Myers SD, Kelly JS, Willems MET. Acute post-exercise effects of concentric and eccentric exercise on glucose clearance. International Journal of Sport Nutrition and Exercise Metabolism 25(1), 14-19, 2015.
Gault ML, Willems, MET. Walking and aerobic capacity in old adults after concentric and eccentric endurance exercise at self-selected intensities. Health 6(8), 654-663, 2014.
Ozmen T, Yuktasir B, Yildirim NU, Yalcin B & Willems MET. Effect of Explosive Strength Training on Speed and Agility Performance in Wheelchair Basketball Athletes. Brazilian Journal of Sports Medicine 20(2), 97-100, 2014.
Gavin JP, Willems MET & Myers SD. Reproducibility of speed of lactate markers during 4 and 8 min stage incremental running: a pilot study. Journal of Science and Medicine in Sport 17(6), 635-639, 2014.
Gault ML & Willems MET. Aging, functional capacity and eccentric exercise training. Aging and Disease, 4(6), 351-363, 2013.
Gault ML & Willems MET. Isometric strength and steadiness adaptations of the knee extensor muscles to level and downhill treadmill walking in older adults. Biogerontology 14(2), 197-208, 2013.
Gavin JP, Myers SD & Willems MET. Physiological responses at various lactate markers for running at 4 and 8 minute treadmill increments. Sport Scientific and Practical Aspects – International Scientific Journal of Kinesiology 10(1), 5-14, 2013.
Blacker SD, Fallowfield JL, Bilzon JLJ, Willems MET. Neuromuscular impairment following backpack load carriage. Journal of Human Kinetics 37, 91-98, 2013.
Blacker SD, Fallowfield JL, Bilzon JLJ, & Willems MET. Intra- and interday reliability of voluntary and electrically stimulated isometric contractions of the quadriceps femoris. Journal of Electromyography and Kinesiology 23(4), 886-891, 2013.
Gault ML, Clements RE & Willems MET. Cardiovascular responses during downhill treadmill walking at self-selected intensity in older adults. Journal of Aging and Physical Activity 21, 335-347, 2013.
Willems MET & Ponte J. Divergent fatigue resistance of dominant and non-dominant quadriceps muscle during voluntary isometric contractions. Journal of Science and Medicine in Sport 16(3), 240-244, 2013.
Willems MET, Sallis CW & Haskell JA. Multi-ingredient supplementation enhances adaptations of resistance training-specific tasks in young males. Journal of Human Kinetics 33, 91-103, 2012.
Gault ML, Clements RE & Willems MET. Functional mobility of older adults after concentric and eccentric endurance exercise. European Journal of Applied Physiology 112(11), 3699-3707, 2012.
Fallowfield JL, Blacker SD, Willems MET, Davy T & Layden J. Neuromuscular and cardiovascular responses of Royal Marine recruits to load carriage in the field. Applied Ergonomics 43(6), 1131-1137, 2012.
Madigan DJ & Willems MET. Effect of caffeine on fatigue during submaximal isometric contractions at different knee angles. Medicina Sportiva 15(4), 194-200, 2011.
Gault ML, Clements RE & Willems MET. Muscle damage from downhill treadmill walking does not change walking economy in healthy older adults. Journal of Human Kinetics 27, 55-65, 2011.
Blacker SD, Williams NC, Fallowfield JL & Willems MET. The effect of a carbohydrate beverage on the physiological responses during prolonged load carriage. European Journal of Applied Physiology 111(8): 1901-1908, 2011.
Webb EC & Willems MET. Effects of wearing graduated compression garment during eccentric exercise. Medicina Sportiva 14 (4): 193-198, 2010.
Blacker SD , Fallowfield JL, Bilzon JLJ & Willems MET. Neuromuscular function following prolonged load carriage on level and downhill gradients. Aviation, Space and Environmental Medicine 81, 745-754, 2010.
Blacker SD, Fallowfield JL, Bilzon JLJ & Willems MET. Within-day and between-days reproducibility of isokinetic parameters of knee, trunk and shoulder movements. Isokinetics and Exercise Science 18(1), 45-55, 2010.
Vatansever-Ozen S, Tiryaki- Sonmez G, Yuktasir B, Yalcin HB, Bugdayci G & Willems M. Effects of exercise on leptin and acylated ghrelin hormones in trained males. Journal of Exercise Physiology online 13(2), 20-30, 2010.
Willems MET, Miller GR, Stauber FD & Stauber WT.. Effects of repeated lengthening contractions on skeletal muscle adaptations in female rats. Journal of Physiological Sciences 60, 143-150, 2010.
Blacker SD , Williams NC, Fallowfield JL, Bilzon JLJ & Willems MET. Carbohydrate vs protein supplementation for recovery of neuromuscular function following prolonged load carriage. Journal of the International Society of Sports Nutrition 2010, 7:2.
Willems MET & Bond TS. Metabolic equivalent of brisk walking and playing new generation active computer games in young-adults. Medicina Sportiva 13(2): 95-98, 2009.
Willems MET & Bond TS. Physiological and metabolic responses to playing Nintendo Wii Sports and brisk treadmill walking. Journal of Human Kinetics 22, 43-49, 2009.
Gault ML, Clements RE & Willems MET. Effect of wearing the Cosmed K4b2 metabolic system on 1 mile walking performance in older adults. Journal of Human Kinetics 21, 41-48, 2009.
Blacker SD, Fallowfield JL, Bilzon JLJ & Willems MET. Physiological responses to load carriage during level and downhill treadmill walking. Medicina Sportiva 13 (2): 116-124, 2009.
Willems MET, Hale T & Wilkinson CS. Effect of massage on delayed onset muscle soreness and performance following downhill walking. Medicina Sportiva 13(2), 61-66, 2009.
Willems MET & Northcott SR. Gender differences for voluntary isometric contractions of knee extensor muscles after downhill running using surface EMG. Medicina Sportiva 13(1): 35-42, 2009.
Willems MET & Stauber WT. The effect of number of lengthening contractions on rat isometric force production at different frequencies of nerve stimulation. Acta Physiologica 196(3): 351-356, 2009.
Willems MET & Stauber WT. Streptomycin and EDTA decrease the number of desmin-negative fibers following stretch injury. Muscle & Nerve 32, 310-315, 2005.
Willems MET & Stauber WT. Attenuation of stretch-induced histopathologic changes of skeletal muscles by quinacrine. Muscle & Nerve 27, 65-71, 2003.
Stauber WT & Willems MET. Prevention of histopathologic changes from 30 repeated stretches of active rat skeletal muscles by long inter-stretch rest times. European Journal of Applied Physiology 88, 94-99, 2002.
Willems MET & Stauber WT. Fatigue and recovery at long and short muscle lengths after eccentric training. Medicine & Science in Sports & Exercise 34, 1738-1743, 2002.
Willems MET & Stauber WT. Effect of contraction history on torque deficits by stretches of active rat skeletal muscle. Canadian Journal of Applied Physiology 27, 323-335, 2002.
Willems MET & Stauber WT. Force deficits by stretches of activated muscles with constant or increasing velocity. Medicine & Science in Sports & Exercise 34, 667-672, 2002.
Willems MET, Miller GR & Stauber WT. Force deficits after stretches of activated rat muscle-tendon complex with reduced collagen cross-links. European Journal of Applied Physiology 85, 405-411, 2001.
Willems MET & Stauber WT. Force deficits after repeated stretches of activated skeletal muscles in female and male rats. Acta Physiologica Scandinavica 172, 63-67, 2001.
Willems MET & Stauber WT. Force during stretches of rat skeletal muscles after hypertonia at short and long lengths. Archives of Physiology and Biochemistry 108, 391-397, 2000.
Willems MET & Stauber WT. Changes in force by repeated stretches of skeletal muscle in young and old female Sprague Dawley rats. Aging Clinical and Experimental Research 12, 478-481, 2000.
Willems MET & Stauber WT. Effect of resistance training on muscle fatigue and recovery in intact rats. Medicine & Science in Sports & Exercise 32, 1887-1893, 2000.
Willems MET & Stauber WT. Performance of plantar flexor muscles with eccentric and isometric contractions in intact rats. Medicine & Science in Sports & Exercise 32, 1293-1299, 2000.
Willems MET & Stauber WT. Force output during and following active stretches of rat plantar flexor muscles: effect of velocity of ankle rotation. Journal of Biomechanics 33, 1035-1038, 2000.
Willems MET, Huijing PA & Fridén J. Swelling of sarcoplasmic reticulum after isometric contractions in rat semimembranosus lateralis muscle. Acta Physiologica Scandinavica 165, 347-356, 1999.
Willems MET & Stauber WT. Isometric and concentric performance of electrically stimulated ankle plantar flexor muscles in intact rat. Experimental Physiology 84, 379-384, 1999.
Willems MET & Purslow PP. Mechanical and structural characteristics of single muscle fibres and fibre groups from raw and cooked pork longissimus muscle. Meat Science 46, 285-301, 1997.
Willems MET & Purslow PP. Effect of postrigor sarcomere length on mechanical and structural characteristics of raw and heat-denaturated single porcine muscle fibres. Journal of Texture Studies 27, 217-233, 1996.
Willems MET & Huijing PA. Hip joint position and architecture of rat semimembranosus muscle: implications for length force characteristics. Acta Anatomica 152, 56-65, 1995.
Willems MET & Huijing PA. Heterogeneity of mean sarcomere length in different fibres: effects on length range of active force production in rat muscle. European Journal of Applied Physiology 68, 489-496, 1994.
Willems MET & Huijing PA. Mechanical and geometrical properties of the rat semimembranosus lateralis muscle during isometric contractions. Journal of Biomechanics 27, 1109-1118, 1994.
Willems MET & Huijing PA. Effect of growth on architecture of rat semimembranosus lateralis muscle. The Anatomical Record 233, 25-31, 1992.
Willems MET, Brozinick Jr JT, Torgan CE, Cortez MY & Ivy JL. Muscle glucose uptake of obese Zucker rats trained at two different intensities. Journal of Applied Physiology 70, 36-42, 1991.
Torgan CE, Brozinick Jr JT, Willems MET & Ivy JL. Substrate utilization during acute exercise in obese Zucker rats. Journal of Applied Physiology 69, 1987-1991, 1990.
Abstracts (2018 only)
MET Willems, M Todaka, MD Cook, Y Sekine. New Zealand blackcurrant powder reduces a human body odor component in middle-aged and older adults. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
MET Willems, MA Sahin, N Parktin, W Widjaja, A Ajjimaporn. Effects of New Zealand blackcurrant extract intake on metabolic and physiological responses during moderate intensity exercise in Caucasian and Thai males. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
BJ Lee, TR Flood, AM Hiles, LEV Wheeler, R Costello, EF Walker, K Ashdown, MR Kuennen, MET Willems. New Zealand Blackcurrant extract enhances fat oxidation during submaximal exercise in the heat. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
BJ Lee, TR Flood, AM Hiles, LEV Wheeler, R Costello, EF Walker, K Ashdown, MET Willems, MR Kuennen. New Zealand Blackcurrant extract attenuates enterocyte damage following exertional heat stress. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
MA Şahin, P Bilgiç, S Montanari, MET Willems.. Effects of duration of New Zealand blackcurrant extract intake on whole-body fat oxidation during moderate intensity exercise in males. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
MA Şahin, P Bilgiç, S Montanari, MET Willems. Effects of daily and intermittent intake of New Zealand blackcurrant extract on whole-body fat oxidation during moderate intensity exercise in males. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
IC Perkins, SD Blacker, MET Willems. Intra-individual responses to New Zealand blackcurrant extract during high-intensity, intermittent running: A repeat response study. International Sport and Exercise Nutrition Conference. Newcastle upon Tyne, UK, 2018.
Mark ET Willems, Tim Berendsen, Mehmet A Şahin, Stephen D Myers, Sam D Blacker, Matthew D Cook. EffecACSM’s 65th Annual meeting, Minneapolis, 2018.
Potter, J.A., Perkins, I.C., Hodgson, C. I., Broadhurst, M., Howells, L., Gilbert, J., Willems, M.E.T. Effect of New Zealand blackcurrant extract on sports climbing performance. ECSS 2018, Dublin, Ireland, 2018.
Shepherd SO, Willems MET, Strauss JA. New Zealand blackcurrant extract enhances fat oxidation during prolonged cycling in endurance-trained females. ECSS 2018, Dublin, Ireland, 2018.
Parktin N, Nana A, Widjaja W, Ajjimaporn A, Willems MET. Effect of New Zealand blackcurrant on cardiovascular responses during sitting rest and treadmill walking in Thai men. ECSS 2018, Dublin, Ireland, 2018.
Willems MET, Parktin N, Widjaja W, Ajjimaporn A. Effect of New Zealand blackcurrant on metabolic and physiological parameters during sitting rest and treadmill walking in Thai men. ECSS 2018, Dublin, Ireland, 2018.
Perkins IC, Bradley M, Blacker SD, Willems MET. Effect of New Zealand blackcurrant extract muscle fibre-specific fatigue. ECSS 2018, Dublin, Ireland, 2018.
Journal Advisory Editorial Board
2009 – present: European Journal of Applied Physiology
Journal Editorial Board
2011 – present: European Journal of Sport Science
2017 – present: Journal of Functional Morphology and Kinesiology
2022 – present: Frontiers in Nutrition (Sport and Exercise Nutrition)